Cytomander the Swift is one of the Spiral King's Four Divine Generals and the commander of Teppelin's aerial forces and owner of the flying battleship, the Dai-Gunten. As General Adiane only gets along with General Thymilph, it seems that Cytomander only gets along with General Guame, just like cytosine and guanine only bond with each other in the double helix of DNA. Cytomander attacked the Great Gurren Brigade twice, losing his life the second time, when the Dai-Gunten exploded. Cytomander's name is based on the DNA chemical cytosine and the elemental Salamander. His Gunmen, Shuzack, is based on the mythical Vermilion Bird, Suzaku.